We may be suffering from Darwin ­fatigue after all those 200th ­anniversary TV programmes last year, but Jon Amiel's sombre film is made with care and delicacy. It transcends costume drama with the hallucinations and ­delusions of the sick scientist (Paul ­Bettany) with an illness that also kills his favourite child. At times, it ­resembles The Singing Detective, also directed by Amiel all those years ago. The nature footage and the illustrated tales he tells the kids are brilliantly pulled off. This adds a layer of meaning to the central, well known science v God dilemma, which troubled Darwin's religious wife.